ε-Globin
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{{#if: P02100 | {{#ifeq: | NV || | style="background:#C3FDB8; color:#000000;" | Homologie-Familie
{{#if: | | style="background:#C3FDB8; color:#000000;" | Übergeordnetes Taxon
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| Struktur von ε-Globin im Homotetramer | |||||||||||||
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| Kofaktor | Häm
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ε-Globin, epsilon-Globin, oder Hämoglobin epsilon-Kette ist ein Protein aus der Familie der Globine, dessen 147 Aminosäuren lange Polypeptidkette ein Häm als Cofaktor bindet und das so Teil des Proteinkomplexes von embryonalen Hämoglobinen ist – bei Hb Gower-1 (ζ2ε2) und bei Hb Gower-2 (α2ε2) –, die Hämoglobin-Untereinheit epsilon (HBE oder HBE1).
Das ε-Globin ist ein Bestandteil von Hämoglobin-Varianten, die in der Embryonalperiode für den Sauerstofftransport gebildet werden. Beim Menschen wird es vom HBB-Gen codiert, das im Gen-Cluster des β-Globin-Locus (5′ – HBE1 – HBG2 – HBG1 – HBD – HBB-3′)<ref>{{#if:Vorlage:Cite book/URL|{{#iferror: {{#iferror:{{#invoke:Vorlage:FormatDate|Execute}}|}}| |}}}}{{#if:{{#if:
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Die Expression dieser einander sehr ähnlichen Gene wird hier über eine gemeinsame Kontrollregion geregelt und ist entwicklungsabhängig unterschiedlich. Je nach Entwicklungsphase bzw. Zellmilieu kann die Proteinbiosynthese von entsprechenden ε-, γ-, δ- oder β-Globinen veranlasst werden.<ref name="pmid2649166">{{#invoke:Vorlage:Literatur|f}}{{#if: | {{#if: Vorlage:Cite book/ParamBool | Vorlage:Toter Link/archivebot | Vorlage:Webarchiv/archiv-bot }} }}{{#invoke:TemplatePar|check |all = title= |opt = vauthors= author= author1= authorlink= author-link= author-link1= author1-link= author2= author3= author4= author5= author6= author7= author8= author9= editor= last= first= last1= first1= last2= first2= last3= first3= last4= first4= last5= first5= last6= first6= last7= first7= last8= first8= last9= first9= last10= first10= last11= first11= last12= first12= last13= first13= last14= first14= last15= first15= others= script-title= trans-title= date= year= volume= issue= number= series= page= pages= at= issn= arxiv= bibcode= doi= pmid= pmc= jstor= oclc= id= url= url-status= format= access-date= archive-date= archive-url= archivebot= offline= location= publisher= language= quote= work= journal= newspaper= magazine= periodical= name-list-style= url-access= doi-access= display-authors= via= s2cid= mr= type= citeseerx= accessdate= archivedate= archiveurl= coauthors= month= day= last16= first16= last17= first17= last18= first18= last19= first19= last20= first20= last21= first21= last22= first22= last23= first23= last24= first24= last25= first25= last26= first26= last27= first27= last28= first28= last29= first29= last30= first30= last31= first31= |cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal |errNS = 0 |template = Vorlage:Cite journal |format = |preview = 1 }}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Blood || Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}{{#if: Vorlage:Cite book/ParamBool | Vorlage:Cite book/Meldung }}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Higgs DR, Vickers MA, Wilkie AO, Pretorius IM, Jarman AP, Weatherall DJ|^^||+1}}{{#ifeq:^^|^^||+1}} > 1 | Vorlage:Cite book/Meldung }}</ref> Diese Proteine unterscheiden sich voneinander nur in wenigen Aminosäuren ihrer jeweils 147 AS langen Kette; doch zeigen die aus ihnen zusammengesetzten Hämoglobine Unterschiede im Sauerstoff-Bindungsvermögen.
Funktion
In der menschlichen Embryogenese wird zunächst das Gen für epsilon-Globin (HBE) aktiviert. Schon während der dritten Entwicklungswoche bilden Zellen in den noch extraembryonal gelegenen Blutinseln des Dottersacks bzw. Nabelschnurbläschens die ersten (embryonalen) Hämoglobine. Diese Proteinkomplexe setzen sich aus vier häm-bindenden Globin-Untereinheiten zusammen, die sich paarweise gleichen. Beim embryonalen Hämoglobin „Hb Gower I“ bestehen die beiden Paare aus epsilon-Kette und zeta-Kette – ζ2ε2); beim „Hb Gower II“ sind es epsilon-Ketten und alpha-Ketten – α2ε2. Beide dieser embryonalen Hämoglobine werden später normalerweise durch fötales Hämoglobin (HbF – α2γ2) und anschließend durch die Hämoglobine Erwachsener (HbA(1) – α2β2; HbA2 – α2δ2) verdrängt.
Einzelnachweise
<references />
Literatur
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| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Proudfoot NJ, Baralle FE|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Nature
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Proudfoot NJ, Brownlee GG|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
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| Vorlage:Toter Link/archivebot
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: J. Biol. Chem.
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Marotta CA, Forget BG, Cohne-Solal M, et al.|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
}}{{#invoke:TemplatePar|check
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Nature
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Gelinas R, Endlich B, Pfeiffer C, et al.|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
}}{{#invoke:TemplatePar|check
|all = title=
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Nature
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
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| Vorlage:Cite book/Meldung
}}
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
}}{{#invoke:TemplatePar|check
|all = title=
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
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|template = Vorlage:Cite journal
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Gene
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}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Lang KM, Spritz RA|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
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| Vorlage:Toter Link/archivebot
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}}{{#invoke:TemplatePar|check
|all = title=
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: J. Clin. Invest.
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Ley TJ, Maloney KA, Gordon JI, Schwartz AL|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
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}}
}}{{#invoke:TemplatePar|check
|all = title=
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
|errNS = 0
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Science
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Chabot B, Black DL, LeMaster DM, Steitz JA|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
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| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
}}{{#invoke:TemplatePar|check
|all = title=
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Proc. Natl. Acad. Sci. U.S.A.
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Engelke DR, Hoener PA, Collins FS|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
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| {{#if: Vorlage:Cite book/ParamBool
| Vorlage:Toter Link/archivebot
| Vorlage:Webarchiv/archiv-bot
}}
}}{{#invoke:TemplatePar|check
|all = title=
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|cat = Wikipedia:Vorlagenfehler/Vorlage:Cite journal
|errNS = 0
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Biochem. Biophys. Res. Commun.
|| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}{{#if: Vorlage:Cite book/ParamBool
| Vorlage:Cite book/Meldung
}}Vorlage:Cite book/Meldung2{{#ifexpr: 0{{#ifeq:^^|^^||+1}}{{#ifeq:^^|^^||+1}}{{#ifeq:Fei YJ, Stoming TA, Efremov GD, et al.|^^||+1}}{{#ifeq:^^|^^||+1}} > 1
| Vorlage:Cite book/Meldung
}}
- {{#invoke:Vorlage:Literatur|f}}{{#if:
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}}{{#invoke:TemplatePar|check
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Proc. Natl. Acad. Sci. U.S.A.
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}}{{#invoke:TemplatePar|check
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Cell
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}}{{#invoke:TemplatePar|check
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: Proc. Natl. Acad. Sci. U.S.A.
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| Vorlage:Toter Link/archivebot
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}}{{#invoke:TemplatePar|check
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}}Vorlage:Cite book/URL{{#if: | Vorlage:Cite book/Meldung }}{{#if: | Vorlage:Cite book/Meldung }}{{#if: J. Biol. Chem.
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Kategorien:
- Wikipedia:Unbekannter Wert (Chemie)
- Wikipedia:Artikel ohne Wikidata-Datenobjekt (Chemie)
- Wikipedia:Proteinbild nicht vorhanden
- Wikipedia:Vorlagenfehler/Parameter:URL
- Wikipedia:Vorlagenfehler/Parameter:Linktext
- Wikipedia:Vorlagenfehler/Parameter:Datum
- Wikipedia:Vorlagenfehler/Vorlage:"
- Wikipedia:Weblink offline fix-attempted
- Wikipedia:Vorlagenfehler/Vorlage:Toter Link
- Wikipedia:Vorlagenfehler/Vorlage:Toter Link/URL fehlt
- Sauerstofftransporter
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