CTD-Phosphatase

{{#if: | {{#if: 1J2X, 1ONV, 2K7L| {{#if:2498| {{#if: 961 Aminosäuren | {{#if: Homodimer | {{#if: TFIIF | {{#if: | {{#if: 3 | {{#if: CTDP1CCFDN, FCP1Q9Y5B06049271926953CTDP1 | {{#if: CTDP1CCFDN, FCP1 | {{#if: Q9Y5B06049271926953CTDP1 | {{#if: | {{#if:| {{#if: | {{#if: 3.1.3.16PhosphataseLösung einer PhosphorsäureesterbindungPhosphoprotein + H₂OProtein + Phosphat | {{#if: 3.1.3.16Phosphatase| {{#if: | {{#if: | {{#if: Lösung einer Phosphorsäureesterbindung| {{#if: Phosphoprotein + H₂O| {{#if: Protein + Phosphat| {{#if: Eukaryoten<ref>InterPro-Eintrag</ref>{{#if: Q9Y5B0 | {{#ifeq: | NV ||1}}|}}| {{#if: Q9Y5B0 | {{#ifeq: | NV || | style="background:#C3FDB8; color:#000000;" | Homologie-Familie {{#if: Eukaryoten<ref>InterPro-Eintrag</ref> | | style="background:#C3FDB8; color:#000000;" | Übergeordnetes Taxon {{#if: | | style="background:#C3FDB8; color:#000000;" | Ausnahmen {{#if: {{#if: Hausmaus | | colspan="3" style="background:#90EE90; color:#202122; text-align:center;" | Orthologe {{#if: ENSG00000060069ENSMUSG00000033323

{{#if: | | CTD-Phosphatase }}
CTD-Phosphatase }}
Andere Namen	{{{Andere Namen}}} }}
Vorhandene Strukturdaten: 1J2X, 1ONV, 2K7L }}
Eigenschaften des menschlichen Proteins }}
Masse/Länge Primärstruktur	961 Aminosäuren }}
Sekundär- bis Quartärstruktur	Homodimer }}
Kofaktor	TFIIF }}
Präkursor	}}
Isoformen	3 }}
Bezeichner
{{#if:CCFDN, FCP1 | Gen-Namen | Gen-Name}} | Gen-Name(n) }}	{{#if:2498 | CTDP1 | CTDP1}}{{#if:CCFDN, FCP1 |, CCFDN, FCP1}} }}
Externe IDs	{{#if:CTDP1 | GeneCards: Vorlage:GeneCards}}{{#if:604927 | OMIM: 604927}}{{#if:Q9Y5B0| {{#if: | | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}}}{{#if:1926953 | MGI: 1926953}}{{#if: | PubChem: {{#switch: {{{PubChem}}}		0 | }}	- = –	valid|1|2=/^[1-9]%d?%d?%d?%d?%d?%d?%d?%d?$/	0 | Wikipedia:Vorlagenfehler/Vorlage:PubChem}}	template= Vorlage:PubChem	format= }} }}{{#invoke:TemplatePar|check	all= 1=	opt= 2=	0 | Wikipedia:Vorlagenfehler/Vorlage:PubChem}}	template= Vorlage:PubChem	format= }}{{#if: {{#invoke:Wikidata|pageId}}||{{#ifeq: 0 | 0 | }} }}}}{{#if: | CAS-Nummer:  }}{{#if: | }} }} }}
Arzneistoffangaben
ATC-Code	}} {{#if:|
DrugBank	[1] }} {{#if:|
Wirkstoffklasse	}} }}
Transporter-Klassifikation
TCDB	[2] {{#if: |
Bezeichnung	}} }}
{{#if:|Inhibitorklassifikation|{{#if:|Enzymklassifikationen|Enzymklassifikation}}}}
EC, Kategorie	{{#if:3.1.3.16| 3.1.3.16}}{{#if: Phosphatase|, Phosphatase}} }}
MEROPS	[3]}}
MEROPS	[4]}}
Reaktionsart	Lösung einer Phosphorsäureesterbindung }}
Substrat	Phosphoprotein + H2O }}
Produkte	Protein + Phosphat }} }}
Vorkommen
{{#if:		[ {{#if:	Hovergen}}]	[http://hogenom.univ-lyon1.fr/query_sequence?seq=Q9Y5B0 {{#if:	Hovergen}}]}} }} }}
Eukaryoten<ref>InterPro-Eintrag</ref> }}
}} }}
	Mensch	Hausmaus
Entrez {{ #if: 9150	9150	}} {{ #if: 67655	67655	}}
style="background:#C3FDB8; color:#202122;" | Ensembl {{ #if: ENSG00000060069	ENSG00000060069	}} {{ #if: ENSMUSG00000033323	ENSMUSG00000033323	}}

}

 |-
 | style="background:#C3FDB8; color:#202122;" | UniProt
 {{ #if: Q9Y5B0
    | | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}
    | |  }}
 {{ #if: Q7TSG2
    | | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q7TSG2}}{{#if:|Vorlage:Abrufdatum}}
    | |  }}
 |-
 {{#if: NM_001202504NM_026295
    | | style="background:#C3FDB8; color:#202122;" | Refseq (mRNA)
      {{ #if: NM_001202504
         | | NM_001202504
         | |  }}
      {{ #if: NM_026295
         | | NM_026295
         | |  }}
    |}}
 |-
 {{#if: NP_001189433NP_080571
    | | style="background:#C3FDB8; color:#202122;" | Refseq (Protein)
      {{ #if: NP_001189433
         | | NP_001189433
         | |  }}
      {{ #if: NP_080571
         | | NP_080571
         | |  }}
    |}}
 |-
 {{#if: 187967980179756623188040795980469695
    | | style="background:#C3FDB8; color:#202122;" | Genlocus
      {{#if: 18 
        | {{#if: 79679801 
            | {{#if: 79756623 
                | |  #if: hg38 |?db=hg38|}}&position=chr18:79679801-79756623 Chr 18: {{#expr: 79679801 / 1000000 round 2}} – {{#expr: 79756623 / 1000000 round 2}} Mb 
                | |  }}
            | |  }}
        | |  }}
      {{#if: 18 
        | {{#if: 80407959 
            | {{#if: 80469695 
                | |  #if: mm10 |?db=mm10|}}&position=chr18:80407959-80469695 Chr 18: {{#expr: 80407959 / 1000000 round 2}} – {{#expr: 80469695 / 1000000 round 2}} Mb 
                | |  }}
           | |  }}
       | |  }}
    |}}
 |-
 | style="background:#C3FDB8; color:#202122;" | PubMed-Suche 
 {{ #if: 9150
    | | 9150
    | |  }}
 {{ #if: 67655
    | | 67655
    | |  }}

| |- | colspan="3" style="background:#90EE90; color:#202122; text-align:center;" | Orthologe (Mensch) |- | style="background:#C3FDB8; color:#202122;" | Entrez | colspan="2" | 9150

 {{#if: ENSG00000060069
    | |-

| style="background:#C3FDB8; color:#202122;" | Ensembl | colspan="2" | ENSG00000060069 }} |- | style="background:#C3FDB8; color:#202122;" | UniProt | colspan="2" | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}

 {{#if: NM_001202504
    | |-

| style="background:#C3FDB8; color:#202122;" | Refseq (mRNA) | colspan="2" | NM_001202504 }}

 {{#if: NP_001189433
    | |-

| style="background:#C3FDB8; color:#202122;" | Refseq (Protein) | colspan="2" | NP_001189433 }}

 {{#if: 18 
   | {{#if: 79679801 
       | {{#if: 79756623 
           | |-

| style="background:#C3FDB8; color:#202122;" | Genlocus | colspan="2" | #if: hg38 |?db=hg38|}}&position=chr18:79679801-79756623 Chr 18: {{#expr: 79679801 / 1000000 round 2}} – {{#expr: 79756623 / 1000000 round 2}} Mb

           |  }}
       |  }}
   |  }}

|- | style="background:#C3FDB8; color:#202122;" | PubMed-Suche | colspan="2" | 9150 }} {{#if: Hausmaus

    | {{#if: 67655ENSMUSG00000033323NM_026295NP_080571mm10188040795980469695Q7TSG2 
        |
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    | {{#if: 67655ENSMUSG00000033323NM_026295NP_080571mm10188040795980469695Q7TSG2 
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        |}}

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 |-
 | style="background:#C3FDB8; color:#202122;" |
 | style="background:#C3FDB8; color:#202122; text-align:center;" | Mensch
 | style="background:#C3FDB8; color:#202122; text-align:center;" | Hausmaus
 |-
 | style="background:#C3FDB8; color:#202122;" | Entrez
 {{ #if: 9150
    | | 9150
    | |  }}
 {{ #if: 67655
    | | 67655
    | |  }}
 |-
 {{#if: ENSG00000060069ENSMUSG00000033323
    | | style="background:#C3FDB8; color:#202122;" | Ensembl
    {{ #if: ENSG00000060069
       || ENSG00000060069
       ||  }}
    {{ #if: ENSMUSG00000033323
       || ENSMUSG00000033323
       ||  }}
  |}}
 |-
 | style="background:#C3FDB8; color:#202122;" | UniProt
 {{ #if: Q9Y5B0
    | | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}
    | |  }}
 {{ #if: Q7TSG2
    | | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q7TSG2}}{{#if:|Vorlage:Abrufdatum}}
    | |  }}
 |-
 {{#if: NM_001202504NM_026295
    | | style="background:#C3FDB8; color:#202122;" | Refseq (mRNA)
      {{ #if: NM_001202504
         | | NM_001202504
         | |  }}
      {{ #if: NM_026295
         | | NM_026295
         | |  }}
    |}}
 |-
 {{#if: NP_001189433NP_080571
    | | style="background:#C3FDB8; color:#202122;" | Refseq (Protein)
      {{ #if: NP_001189433
         | | NP_001189433
         | |  }}
      {{ #if: NP_080571
         | | NP_080571
         | |  }}
    |}}
 |-
 {{#if: 187967980179756623188040795980469695
    | | style="background:#C3FDB8; color:#202122;" | Genlocus
      {{#if: 18 
        | {{#if: 79679801 
            | {{#if: 79756623 
                | |  #if: hg38 |?db=hg38|}}&position=chr18:79679801-79756623 Chr 18: {{#expr: 79679801 / 1000000 round 2}} – {{#expr: 79756623 / 1000000 round 2}} Mb 
                | |  }}
            | |  }}
        | |  }}
      {{#if: 18 
        | {{#if: 80407959 
            | {{#if: 80469695 
                | |  #if: mm10 |?db=mm10|}}&position=chr18:80407959-80469695 Chr 18: {{#expr: 80407959 / 1000000 round 2}} – {{#expr: 80469695 / 1000000 round 2}} Mb 
                | |  }}
           | |  }}
       | |  }}
    |}}
 |-
 | style="background:#C3FDB8; color:#202122;" | PubMed-Suche 
 {{ #if: 9150
    | | 9150
    | |  }}
 {{ #if: 67655
    | | 67655
    | |  }}

| |- | colspan="3" style="background:#90EE90; color:#202122; text-align:center;" | Orthologe (Mensch) |- | style="background:#C3FDB8; color:#202122;" | Entrez | colspan="2" | 9150

 {{#if: ENSG00000060069
    | |-

| style="background:#C3FDB8; color:#202122;" | Ensembl | colspan="2" | ENSG00000060069 }} |- | style="background:#C3FDB8; color:#202122;" | UniProt | colspan="2" | {{#if: kurz| | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}

 {{#if: NM_001202504
    | |-

| style="background:#C3FDB8; color:#202122;" | Refseq (mRNA) | colspan="2" | NM_001202504 }}

 {{#if: NP_001189433
    | |-

| style="background:#C3FDB8; color:#202122;" | Refseq (Protein) | colspan="2" | NP_001189433 }}

 {{#if: 18 
   | {{#if: 79679801 
       | {{#if: 79756623 
           | |-

| style="background:#C3FDB8; color:#202122;" | Genlocus | colspan="2" | #if: hg38 |?db=hg38|}}&position=chr18:79679801-79756623 Chr 18: {{#expr: 79679801 / 1000000 round 2}} – {{#expr: 79756623 / 1000000 round 2}} Mb

           |  }}
       |  }}
   |  }}

|- | style="background:#C3FDB8; color:#202122;" | PubMed-Suche | colspan="2" | 9150 }} {{#if: Hausmaus

    | {{#if: 67655ENSMUSG00000033323NM_026295NP_080571mm10188040795980469695Q7TSG2 
        |
        |  Spezies2= ist definiert, es sind jedoch keine weiteren Angaben zu Spezies2 vorhanden.{{#ifeq:0|0| }} }}
    | {{#if: 67655ENSMUSG00000033323NM_026295NP_080571mm10188040795980469695Q7TSG2 
        |  Es sind Angaben zu Spezies2 vorhanden, Spezies2= ist jedoch nicht definiert.{{#ifeq:0|0| }}
        |}}

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|}{{#if:Protein_CTDP1_PDB_1onv.png||}}

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Die CTD-Phosphatase (FCP1) (genauer TFIIF-assoziierende CTD-Phosphatase) ist das Enzym, das in Eukaryoten während der Transkription die Phosphatreste von Serin-2 und Serin-5 der Polymerase II entfernt, womit die Polymerase für die Elongation (re-)aktiviert wird. Es handelt sich also um eine Phosphatase. Das Enzym ist beim Menschen in allen Gewebetypen zu finden. Mutationen im entsprechenden CTDP1-Gen können zu erblichen multiplen Entwicklungsstörungen mit Katarakt, dem sogenannten CCFDN-Syndrom führen.<ref>{{#if: | | UniProt }} {{#if:|{{{titel}}}|Q9Y5B0}}{{#if:|Vorlage:Abrufdatum}}</ref>

Um ihre volle Aktivität zu erhalten, wird die CTD-Phosphatase selbst von Caseinkinase 2 phosphoryliert, und benötigt die RAP74-Untereinheit von TFIIF als Cofaktor.<ref>{{#invoke:Vorlage:Literatur|f}}{{#if:

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</ref>

Die Phosphorylierung von FCP1 wird von den Proteinen Pin1 und Hce1 sowie vom Tat-Protein des HI-Virus gehemmt. FCP1 wird von der Arginin-Methyltransferase PRMT5 methyliert und ein FCP1-PRMT5-Komplex methyliert Histon H4.<ref>{{#invoke:Vorlage:Literatur|f}}{{#if:

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     | Vorlage:Cite book/Meldung
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     | Vorlage:Cite book/Meldung
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Weblinks

• {{#ifeq:|ID|48431|

Eintrag zu {{#if:Kongenitale Katarakt-Gesichtsdysmorphie-Neuropathie-Syndrom|Kongenitale Katarakt-Gesichtsdysmorphie-Neuropathie-Syndrom|CTD-Phosphatase}}. In: Orphanet (Datenbank für seltene Krankheiten)Vorlage:Abrufdatum }}{{#invoke:TemplatePar|valid|ID|N>0 |cat= {{#ifeq: 0 | 0 | Wikipedia:Vorlagenfehler/Orphanet}} |template= Vorlage:Orphanet }}{{#invoke:TemplatePar|check |all= ID= Name= |opt= Abruf= Format= |cat= {{#ifeq: 0 | 0 | Wikipedia:Vorlagenfehler/Orphanet}} |template= Vorlage:Orphanet }}

• Gopinathrao / reactome: Hypophosphorylation of RNA Pol II CTD by FCP1P protein

Einzelnachweise

<references />